Fig. 12From: Hydroxylases involved in terpenoid biosynthesis: a reviewProposed partial biosynthetic pathway of carnosic acid, carnosol and 11,20-dihydroxy sugiol. FSs: ferruginol synthases, including CYP76AH1, CYP76AH3, CYP76AH4–7, and CYP76AH22–24. HFSs: 11-hydroxyferruginol synthasesBack to article page